Albert Lasker basic Medical Research Award laureate, Nobel Prize winner, and geneticist Har Gobind Khorana passed away in 2011 at the age of 89. Born in Punjab, British India in 1922, Khorana was raised and schooled in the area before receiving his BA from Punjab University in 1943. He would later gain his MS in 1945 and his PhD from Liverpool University in 1948. Khorana then moved to the United States as a postdoctoral fellow at the University of British Columbia, and was eventually appointed to the faculty of the University of Wisconsin-Madison in 1960. Khorana's groundbreaking research on the biological synthesis of macromolecules, specifically nucleotides, garnered him numerous awards and honors over the course of his career. These included the Albert Lasker Basic Medical Research Award in 1976, the National Medal of Science in 1987, the Louisa Gross Horwitz Prize in 1989 and the Nobel Prize in Physiology or Medicine in 1968. He is survived by his three children, Julia, Emily, and Dave.